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PhD Degree (23)-Fully Funded at University of Copenhagen, Denmark

University of Copenhagen, Denmark invites online Application for number of  Fully Funded PhD Degree at various Departments. We are providing a list of Fully Funded PhD Programs available at University of Copenhagen, Denmark.

Eligible candidate may Apply as soon as possible.

 

(01) PhD Degree – Fully Funded

PhD position summary/title: PhD scholarship in Grundtvig Studies at the Faculty of Theology

WWD investigates how the influential Danish theologian, thinker, and poet N.F.S. Grundtvig worked with textual sources from antiquity to his own day. In addition to the analysis of Grundtvig’s published works, the project group will take advantage of a new research infrastructure called Grundtvig’s Manuscripts Online (GMO). GMO uses the AI-software Transkribus to transcribe of more than 80,000 unprinted pages in the Grundtvig Archive at the Royal Danish Library. The shared theoretical foundations for the interdisciplinary research team are Modern Genre Studies and Conceptual History. 

Deadline : October, 1 , 2026

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(02) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in virus-host interactions at the Department of Immunology and Microbiology

Our core research interest is how viruses hijack cellular systems. A particular focus on this relates to RNA biology. RNA viruses are responsible for significant morbidity and mortality worldwide. Pathogens of our interest include hepatitis viruses, flaviviruses (e.g. tick-borne encephalitis, yellow fever, and zika virus), alphaviruses (e.g. sindbis, chikungunya) and animal pathogens such as pestiviruses. We have specific interest in virus-host interactions, including RNA interactions that are becoming increasingly important, also as potential therapeutic targets. We recently identified and characterized interactions for several viruses. Much, however, remains to be understood on virus-host interactions during infection, including implications for innate immunity. Recent publications from our group include:

  • Larsen, Sherwood, Rivera-Rangel et al. 2026 Nat Comms: “Discovery of 5’ NAD capped viral RNAs reveals evolutionary divergent 5’ metabolite capping across hepaciviruses”
  • Damas, Seemann, Costa, Krambrich et al. 2026 PLoS Pathog: “Characterization of the virus-host RNA-RNA interactome across important human pathogenic RNA viruses” (PMID: 42139316)
  • Sherwood, Rivera-Rangel et al. 2023 Nature: “Hepatitis C virus RNA is 5′-capped with flavin adenine dinucleotide” (PMID: 37407817)
  • Fossat et al. 2023 Cell Rep: “Identification of the viral and cellular microRNA interactomes during SARS-CoV-2 infection” (PMID: 36961814)
  • Wolfisberg et al. 2022 Hepatology: “Neutralization and receptor use of infectious culture-derived rat hepacivirus as a model for HCV” (PMID: 35445423)

Your project on virus-host interactions will be defined together with your supervisor, Troels Scheel, and will involve methodology such as virus cultivation, molecular biology assays, functional characterization, translation and replication reporter assays, crosslinking immune-precipitation (CLIP), RNA-seq and related methods. You will be expected to actively participate in method development and development of the project. A high level of motivation, ambition and involvement will be expected.

Deadline : 31 August 2026

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(03) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in collection-based Natural History Research

The Natural History Museum Denmark offers 1-2 fully funded PhD positions within Collection-based Natural History Research. The successful candidate will conduct 3 years of collection-based research. The PhD project is part of the Museums strategic aim to expand research on unknown species and taxonomic groups that are not well defined and described to provide crucial natural history understanding, data and expertise for developing solutions to planetary challenges, including climate change and biodiversity loss. Potential projects could include but are not limited to dark taxa and unknown species, preferably including representatives of the flora, fauna or geology of Denmark, Greenland or the Faroe Islands.

Deadline : August 30, 2026

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(04) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowships in Cardiac (patho-)Physiology at the Department of Biomedical Sciences

We are looking for a dedicated PhD student, who will be working on understanding mechanism of cardiac pathophysiology. The PhD student(s) recruited for this project will work with several of the many translational modalities in the Cardiac Physiology, spanning from long-term porcine and rodent models of cardiac diseases, ex vivo (human) tissue analyses, electrophysiology, biochemistry, histology, and cMRi, to disclose the underlying mechanisms of cardiac disease from molecule to beating hearts.

Deadline : 30 August 2026

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(05) PhD Degree – Fully Funded

PhD position summary/title: PhD scholarship in the project ‘A Danish model for individual Cognitive Stimulation Therapy for people with dementia’ at the Department of Psychology, University of Copenhagen (UCPH)

The PhD student will be expected to contribute to all parts of the study, including development of the manual and intervention methods and -tools (based on user-involvement and collaboration with stakeholders), recruiting participants, collecting and analysing data, and disseminating findings.  The PhD student will be responsible for collecting and analysing data in particular.

The successful candidate will be working in close cooperation with the PI as well as a student assistant, who will assist with a variety of tasks related to the project. The PhD student will benefit from being part of the Copenhagen Neuropsychology Lab that is part of the research section Cognition and Neuropsychology, where the PhD will partake in common research meetings and daily interaction with other PhD students and researchers. An international stay is planned as part of the PhD.

Deadline : 21 August 2026

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(06) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in greenhouse gases, forest soils and biogeochemistry

The PhD position aims to fill this knowledge gap by addressing how tree species, soil types and forest management practices impact net fluxes of greenhouse gases from the forest soil, focusing primarily on the strong greenhouse gas N2O, but also including CH4 and CO2. The project involves working with existing unpublished datasets from Danish forests, fieldwork at selected forest sites in Denmark with different tree species (conifers and broadleaves) and land use legacies (agriculture and forest), and doing laboratory mesocosm experiments. You will be measuring greenhouse gas fluxes using advanced methods for gas detection in the field and laboratory, including stable isotope techniques, and deploy mathematical and statistical tools to analyze the data.

Deadline : 16 August 2026

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(07) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in geochronology and mineralogy

The main task of the Ph.D. is to characterise heavy mineral populations, including the U-Pb ages of zircons in both metasedimentary source rocks as well as in modern detrital sediments. The main supervisor is Professor Tod Waight.

Deadline :  16 August 2026

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(08) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in molecular nanomaterials

We develop new organic conjugated molecules and nano materials made from these. This includes fluorescent and electroactive molecules  with potential applications in imaging, sensing and energy materials.    

Deadline : August 16 2026.

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(09) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in pharmaceutical physical chemistry

The PhD project focuses on the molecular mechanisms by which small-molecule permeation enhancers increase the absorption of peptide-based therapeutics across the gastrointestinal epithelium. Oral peptide delivery is one of the major challenges in pharmaceutical science, and the chemical and biophysical basis of permeation enhancement remains poorly understood. The aim of this project is to combine biophysical methods, model membrane systems, epithelial permeation assays, and advanced microscopy to provide new molecular insight into oral peptide delivery mediated by penetration enhancers and to support the rational design of improved permeation enhancers candidates.

The PhD candidate will investigate how permeation enhancers interact with model membranes, how they alter membrane structure and dynamics, and how these changes may facilitate the translocation of peptide drugs. This molecular-level work will be complemented by epithelial permeation and other cell-based assays, as well as by confocal microscopy, linking information from simplified model systems to behaviour in cellular systems. The project combines a range of experimental biophysical methods, including lab-based techniques and experiments at large-scale neutron and X-ray scattering facilities in Europe.

Deadline : 16 August 2026

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(10) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in Research Integrity

The project concerns non-financial conflicts of interest in research and whether these should be declared (and if so how). Strongly held theoretical, ethical, political, or religious views can bias researchers in ways similar to financial ties to external organizations. Since the latter must be declared in research publications as potential financial conflicts of interest, it is common to apply the same logic to the former and expect researchers to also declare potential non-financial conflicts of interest in research publications. The Danish Code of Conduct for Research Integrity, for instance, states that:

“Responsible conduct of research includes disclosure of all potential conflicts of interest. This allows financial or other interests to be assessed on an informed basis to evaluate potential bias of the researcher’s professional judgement” (p. 21)

However, there are still many open questions regarding declarations of potential non-financial conflicts of interest. This call invites a PhD fellow to dive into some of these questions using social science methods. Relevant questions include, but are not limited to, how such declarations are used by readers of scientific papers and popular communication of scientific results, how use depends on the status of the sender (e.g., the scientist, the scientific discipline, the university, and the journal), and whether there are specific formats for disclosing nonfinancial conflicts of interest that are particularly useful.

Applicants are encouraged to submit a brief (maximum 3 pages) non-binding research proposal to illustrate their general interest in the topic and ability to formulate a research idea. The final research topic and methods will be determined in dialogue with the supervisors.        

Deadline :16th of August 2026

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(11) PhD Degree – Fully Funded

PhD position summary/title: PhD position in Time-Resolved Crystallography of Enzymes (STRIDE project) — synchrotron-based studies of esterases at MicroMAX

STRIDE aims to develop new methods for time-resolved serial-synchrotron crystallography (TR-SSX) at the MicroMAX beamline (MAX IV Laboratory) to study esterases of the α/β hydrolase superfamily, a major enzyme class with applications in industrial biotechnology and drug development. The main model system is bacterial glucuronoyl esterases, which cleave ester bonds linking lignin and carbohydrates in plant biomass, and have been exensively studied by the project group.

The PhD student will be based at UCPH and supervised by Prof. Leila Lo Leggio. The project has two closely connected strands: 1) synthesis and evaluation and of novel substrate analogues for esterases, including slowly cleavable and photoswitchable compounds; and 2) static single-crystal and serial-synchrotron crystallography of these enzymes in complex with the new analogues, progressing to time-resolved SSX experiments at MicroMAX. The work will involve close collaboration with coworkers at UCPH and secondments to the University of Gothenburg/Chalmers University of Tcehnology.

The student will gain experience across the structural enzymology pipeline, from substrate design to cutting-edge time-resolved crystallography, and will contribute to dissemination activities.

Deadline : 15 August 2026,

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(12) PhD Degree – Fully Funded

PhD position summary/title: The Faculty of Law at the University of Copenhagen announces a number of fully funded PhD positions to commence in February 2027

The goal of the PhD programme is threefold:

(1) to educate a new generation of scholars to produce high-quality independent research;

(2) to equip PhD candidates with analytical and problem-solving skills required for a successful career in academia or in other positions in both the public and private sectors.   

(3) to create societal impact through the production of new knowledge and research skills.

The Faculty of Law offers supervision by highly qualified academics and provides opportunity to research contemporary legal issues in an intellectually stimulating environment. 

The call is open to applicants from across the field of legal scholarship, and the Faculty of Law has a particular interest in proposals for PhD projects that fall within the research areas encompassed by the obligatory courses of the Danish Bachelor of Laws (LL.B). These include, in particular, Company law, Property and Creditor Law, Law of Obligations, Family Law and Law of Wills and Succession, Legal history, Criminal Law and Criminal Procedure, as well as the BA-elective course Labour Law;

Deadline :13 August 2026

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(13) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in Metabolic Control of Glucose Utilization and Homeostasis

Maintenance of energy and glucose homeostasis is essential for metabolic health. Our group has previously shown that glucose utilization in skeletal muscle and liver is governed by coordinated regulation through reversible phosphorylation and allosteric control, using novel knock-in mouse models expressing metabolic enzymes carrying phospho-deficient or allosteric modulator-binding-deficient mutations. This project will focus on defining how glucose phosphorylation by hexokinase 2, together with its allosteric negative feedback regulation, controls skeletal muscle glucose flux and utilization in vivo, and how this contributes to whole-body glucose homeostasis under both physiological and pathophysiological conditions. The successful candidate will perform biochemical analyses combined with structure-guided mutagenesis (cell-free and cell-based assays) to identify allosteric modulator-resistant mutants, as well as in vivo preclinical studies in multiple genetic mouse models, including metabolic phenotyping and assessment of glucose homeostasis.

Deadline :August 10, 2026,

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(14) PhD Degree – Fully Funded

PhD position summary/title: Research Assistent and PhD in Precision Medicine from Graph-Based Medical Digital Twins

The PhD project focuses on developing AI-based approaches to better understand and predict cardiometabolic disease using large-scale health and biological data. The work will contribute to building medical digital twins: computational representations of individuals that capture health trajectories over time. By combining diverse data sources and modern machine learning methods, the project aims to improve how we model disease risk, progression, and treatment response. The research sits at the intersection of data science, biology, and medicine, and involves close collaboration with clinical and international partners. The position offers a unique opportunity to work with world-leading datasets and contribute to the development of next-generation precision health approaches.

Deadline : August 10, 2026

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(15) PhD Degree – Fully Funded

PhD position summary/title: Research Assistant and PhD fellow in the Genetic architecture of skeletal muscle

The overall aim of this PhD project is to leverage MRI data from the UK Biobank to study skeletal muscle traits at scale and to identify genetic factors influencing variation in skeletal muscle composition.

The project will:

  • Generate robust imaging‑derived measures related to skeletal muscle using population‑based MRI data.
  • Identify genetic variants associated with these skeletal muscle traits.
  • Explore the relevance of genetic findings for metabolic health and disease risk using genetic and epidemiological approaches.

Deadline : August 10, 2026,

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(16) PhD Degree – Fully Funded

PhD position summary/title: Research Assistant and PhD fellow in Energy Metabolism

The project focuses on understanding the causal mechanisms underlying the progression of obesity using rodent models and approaches from neuroscience, endocrinology and metabolic physiology. The work will be diverse and include rodent experiments, ex vivo analyses and bioinformatic approaches to investigate how biological systems regulating energy balance adapt during obesity development.

Deadline : August 10, 2026

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(17) PhD Degree – Fully Funded

PhD position summary/title: Research Assistant and PhD fellow in Molecular Nutrition and Cardiometabolic Diseases

The successful candidate will join an interdisciplinary and international research group. The PhD project will focus on understanding how sustainable dietary patterns influence cardiometabolic diseases. The project specifically aims to uncover molecular mechanisms linking plant-based diets to cardiometabolic health by integrating high-dimensional multi-omics data, including metabolomics, proteomics, and genetics, with detailed dietary, clinical, and epidemiological data from large population-based cohorts and a randomized controlled trial (PLANETDIET (read here).

The candidate will work with high-throughput omics data and apply advanced statistical and computational approaches to identify diet-related molecular signatures and investigate their role in the development of cardiometabolic outcomes. The project includes both observational and interventional components, providing a unique opportunity to study causal pathways and validate findings across diverse study designs and international cohorts.

The specific work tasks will include management and harmonization of large-scale datasets, statistical and bioinformatic analyses of multi-omics data, and their integration with dietary and clinical variables. Additional responsibilities include writing and publishing scientific papers, presenting findings at international conferences, and contributing to the center and university activities. The candidate will be encouraged and supported to develop independent research ideas within the scope of the project. Opportunities for collaboration with international partners, including potential research stays abroad, will be encouraged. Teaching and supervision opportunities at the undergraduate or graduate level may also be offered. The PhD candidate will need to complete PhD courses or other equivalent education corresponding to approximately 30 ECTS points.

Deadline : August 10, 2026

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(18) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in solid state Pharmacy at the Department of Pharmacy

Poor aqueous solubility is one of the greatest unsolved challenges in modern drug development, affecting more than 90% of small-molecule drug candidates and severely limiting their therapeutic potential. While crystalline drug forms provide physical stability, their poor solubility often compromises bioavailability and clinical performance. Amorphous drug forms can overcome this limitation by dramatically enhancing solubility; however, their inherent instability and tendency to recrystallize remain major barriers to their widespread application.

This project seeks to challenge conventional paradigms in pharmaceutical formulation by pioneering new concepts in “amorphous engineering”. Recent breakthroughs from our group have demonstrated that drugs can exist in multiple amorphous forms and/or polyamorphs, with distinct physicochemical properties. This largely unexplored phenomenon, known as amorphous variability, opens unprecedented opportunities for the design of next-generation medicines.

The project will investigate the formation, stabilization, and confinement of amorphous forms of drugs within biocompatible particles, creating innovative formulations capable of combining enhanced solubility with improved physical stability, without the need for additional excipients. By integrating state-of-the-art experimental characterization techniques with computational modelling, this research aims to establish transformative strategies for poorly water-soluble drugs and accelerate the development of more effective medicines for patients.

Deadline : 7 August 2026

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(19) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in Gut Microbiome and Cardiometabolic Health

MHI is an initiative that aims to show that the human gut microbiome plays a causal role in cardiometabolic health. As part of this, several human intervention studies are to be conducted testing the effects of microbial species or microbial metabolites on outcomes relevant for cardiometabolic health.

The envisioned PhD project will investigate whether rectal administration of a microbiota-derived metabolite affects colonic hormone secretion and metabolism in humans. A placebo-controlled cross-over trial in human participants will be conducted at Steno Diabetes Center Copenhagen and combine measurements of gut hormones and human metabolism and state-of-the-art omics technologies on biological samples (i.e. transcriptomics, microbiomics and metabolomics). In addition, the PhD student will contribute to other ongoing related research projects.

In your role as PhD student, you will be in charge of writing the study protocol for approval to the Ethical Committee. Following approval, you will be responsible for the recruitment of study participants and for conducting the cross-over intervention study in collaboration with other staff and students. This includes conducting examination visits, handling biological samples, and potentially taking rectal biopsies. Furthermore, you will conduct liquid chromatography mass spectrometry-based metabolomics on biological samples from this and other trials, and analyze transcriptomics and microbiome data obtained from the studies. Finally, you will integrate the clinical intervention-responses with obtained omics data and write scientific publications.

Deadline : 2 August 2026

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(20) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowships in Computational Quantum Physics (Theory & Simulations)

In the Theory and Simulation (T&S) team, we develop the computational methods and capabilities nessecary to realize hardware for practical quantum computing. For this purpose, we employ multiscale modelling and simulate at scales ranging from the atomic to the full device scale. In our work, we are collaboring closely with all of the different teams at NQCP, see https://nqcp.ku.dk/.

We are looking for motivated PhD candiates to undertake research within computational quantum physics. We are interested in developing a project plan with prospective candidates to best match their skills and experience to the activities in the T&S team. We imagine that the candidates would undertake research within one or more of the following topics:

  1. Material simulations
  2. Designs of advanced quantum chips
  3. Hybrid quantum devices
  4. Quantum error correction

Deadline : 1 August 2026

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(21) PhD Degree – Fully Funded

PhD position summary/title: PhD Fellow in Plant Protein Nanofibrils

Protein nanofibrils (PNF) are uniquely suited as functional structuring materials due to their highly ordered molecular architectures. The diverse compositions of plant proteins form different types of PNFs. The relation between these multiscale architectures and morphological differences remain unclear. Therefore, we will investigate the mechanical properties of the different PNF-types at the nanoscale level. The aim is to develop plant PNFs as functional building blocks to improve food texture. The PhD will employ a mix of protein chemistry, microscopic analysis, and quality evaluation to study protein variability, fibrillization methods, interplay between PNF structure, functionality and gel texture.

Deadline : 28 July 2026

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(22) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship in Radiopharmaceutical Chemistry at the Department of Drug Design and Pharmacology

Glioblastoma remains one of the most challenging cancers to treat due to its aggressive nature and the limited efficacy of current therapeutic approaches. Pretargeted radioimmunotherapy offers a promising strategy to improve tumor targeting while reducing radiation exposure to healthy tissues.

This PhD project will focus on the development of novel pretargeted radiopharmaceuticals for imaging and therapy of glioblastoma by targeting the urokinase plasminogen activator receptor (uPAR). The project will utilize bioorthogonal tetrazine ligation chemistry to enable selective in vivo targeting and will investigate the use of fluorine-18 for imaging as well as iodine-131 and astatine-211 for targeted radionuclide therapy.

The successful candidate will design and synthesize new tetrazine-based radioligands with improved stability, including the development of novel tetrazine scaffolds resistant to deastatination. The project will further involve radiolabeling, characterization, and optimization of these compounds, as well as exploration of strategies to improve target-to-background ratios. In parallel, site-specific conjugation approaches will be developed for anti-uPAR antibodies to generate highly defined pretargeting agents.

The PhD student will work closely with an interdisciplinary team within the PreBreach project, combining expertise in radiochemistry, protein engineering, biology, and molecular imaging. Biological evaluation and in vivo studies will be performed in collaboration with project partners, providing a unique opportunity to contribute to the development of next-generation radiopharmaceutical therapies for brain cancer.

Deadline : 24.07.2023

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(23) PhD Degree – Fully Funded

PhD position summary/title: PhD fellowship at Department of Neuroscience

Your tasks will be performing in vitro patch clamp experiments in mouse brain slices; making microsurgeries to implant chronic cranial windows and novel brain stimulation probes in transgenic mice, training the mice, and performing two-photon imaging in the awake mice in a longitudinal study. Besides, you will be involved in data analysis.

Your key tasks as a PhD student at SUND are:

  • Carrying through an independent research project under supervision. 
  • Completing PhD courses or other equivalent education corresponding to approximately 30 ECTS points.
  • Participating in active research environments including a stay at another research team.
  • Obtaining experience with teaching or other types of dissemination related to your PhD project
  • Teaching and disseminating your knowledge.
  • Writing a PhD thesis on the grounds of your project

Deadline : 23th / July 2026

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About The University of Copenhagen, Denmark – Official Website

The University of Copenhagen is a public research university in Copenhagen, Denmark. Founded in 1479, the University of Copenhagen is the second-oldest university in Scandinavia, and ranks as one of the top universities in the Nordic countries and Europe.

Its establishment sanctioned by Pope Sixtus IV, the University of Copenhagen was founded by Christian I of Denmark as a Catholic teaching institution with a predominantly theological focus. In 1537, it was re-established by King Christian III as part of the Lutheran Reformation. Up until the 18th century, the university was primarily concerned with educating clergymen. Through various reforms in the 18th and 19th century, the University of Copenhagen was transformed into a modern, secular university, with science and the humanities replacing theology as the main subjects studied and taught.

The University of Copenhagen consists of six different faculties, with teaching taking place in its four distinct campuses, all situated in Copenhagen. The university operates 36 different departments and 122 separate research centres in Copenhagen, as well as a number of museums and botanical gardens in and outside the Danish capital. The University of Copenhagen also owns and operates multiple research stations around Denmark, with two additional ones located in Greenland. Additionally, The Faculty of Health and Medical Sciences and the public hospitals of the Capital and Zealand Region of Denmark constitute the conglomerate Copenhagen University Hospital.

A number of prominent scientific theories and schools of thought are namesakes of the University of Copenhagen. The famous Copenhagen Interpretation of quantum mechanics was conceived at the Niels Bohr Institute, which is part of the university. The Department of Political Science birthed the Copenhage

 

 

 

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